What are the themes in the story Tuck Everlasting?

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One additional theme is that of civilization vs. nature. The Foster family owns the forest, but because they're immortal they are never truly a part of it. The world of nature is subject to change, decay, and death. As the Fosters exist outside that realm they can never fully understand the significance of the forest. Fortunately, nature proves itself a good deal more effective at protecting itself than the Fosters are able to do. (Cows walk around, rather than through the forest, for example, thus avoiding the spring.) This is not a surprise; nature, because it is subject to change, can adapt...

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One of the main themes of Tuck Everslasting is that death is a natural and necessary part of life.  Throughout history, humans have searched and longed for a cure to death. Tales of a "fountain of youth" have been recounted for thousands of years.  In Tuck Everlasting, this folklore is a reality.  Though the Tuck family have indeed conquered death and become immortal, the book questions whether or not this is necessarily a good thing.

Winnie and Mile's conversation while fishing is one of the most important scenes in the book, and it centers on that exact question. When Winnie says it would be nice if nothing ever had to die, Miles points out that if so, there'd be so many creatures on the earth, they'd be "squeezed in right up next to each other before long." Miles here is trying to show Winnie that death is natural. When the fish he catches is about to die and Winnie protests, he tells her killing things is the natural way.  They let the fish go, but Miles lets her know that it cannot always be that way. 

Winnie learns that the Tucks are not only protecting the secret of their immortality for their own sakes, but for the sake of mankind.  The love the Tuck family shows Winnie is necessary for Winnie to learn this valuable lesson, which she does. In the end, she chooses life AND death.  In the epilogue, Tuck's comment, "good girl," when looking at Winnie's grave, confirms his belief that death is the natural way of the world.  Even though he is grief-stricken, he is happy she made the right choice.