In "Julie of the Wolves", what do the wolf signals mean - biting, nose, ears?

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When the wolves want to express their admiration and homage towards their leader, they bite him gently under the chin.  To signal his approval in response to their expression of love, the leader wags his tail and bites the top of the underling's nose.  Miyax first witnesses this behavior when Nails and Silver approach Amaroq in this manner.  Nails spanks the ground with his forepaws and carefully takes Amaroq'a jaw in his mouth, and "the leader...

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