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The best way to respond to this section of the article might be to say that neither nature nor nuture is completely responsible for the development of human temperament. Both factors play important roles. On the one hand, children with the same genotype developed very differently depending on the environment in their childhood homes. This would tend to point to the importance of "nurture." On the other, there was also a significantly higher risk of children with a certain genotype to respond negatively to such an environment than children who did not have this genotype. Some genetic variants, particularly those that regulate seratonin levels, could have a marked effect on how children respond to negative environments.
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